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It's certainly an interesting‐looking game, and the honey/egg mechanics very on‐theme. Has an element of win‐by‐marriage too which I haven't seen in a while.

Thank you a lot. I have followed your suggestions and made the necessary changes. 

A Colony becomes suffocated when the ratio of pieces vs empty cells is more than 2:1

Counted within that colony (the 1^st 6 ranks), I assume? That's a lot of bookkeeping for over‐the‐board play, though of course computed implementation would make this (and the randomisation of hatchlings) trivial

That's very simple, actually. It's not that hard to count 19 empty cells on the first, or last, six ranks. 

Please let me know if there is anything else to consider.